Research on primate, rodent, antelope, pig, and bird biodiversity, taxonomy, biogeography, behavior, and ecology, mainly in Botswana, Equatorial Guinea, Uganda, Tanzania, Kenya, and Saudi Arabia since 1970. Led research and conservation programs and projects for New Zoological Society, WWF-US, WWF-International, Zoo Atlanta, Conservation International, Arcadia University, Drexel University, Zoological Society of London, and Lolldaiga Hills Ranch. Provided about 100 species and subspecies degree of threat assessments to the IUCN Red List. Mentored and trained many students and field workers, particularly in Uganda, Kenya, and Equatorial Guinea.
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